Publications by theme

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List of publications 

44Leclerc, C.*, F. Courchamp & C. Bellard, Future climate change vulnerability of insular mammals, proofs Nature Communications.

43  Lenzner B., Latombe G., Capinha C., Bellard C., Courchamp F., Diagne C., Dullinger S., Golivets M., Severin D.H. Irl, Kuhn I., Leung B., Liu C., Moser D., Roura-Pascual N., Seebens H., Turbelin A., Weigelt P., Essl F. (2020) What will the future bring for biological invasions on islands? An expert-based assessment, 8, 280

42– Jaric, I.; Bellard, C.; Correia, R.; Courchamp, F.; Douda, K.; Essl, F.; Jeschke, J.; Kalinkat, G.; Kalous, L.; Lennox, R.; Novoa, A.; Proulx, R.; Pyšek, P.; Soriano-Redondo, A.; S., A.; Vardi, R.; Verissimo, D.; Roll, U. Invasion culturomics and iEcology approaches to better understand biological invasions, proofs in Conservation Biology

41Bellard, C.; Hugueny, B Importance of metapopulation dynamic to explain fish persistence in a river system, in Freshwater Biology. 2020; 00: 1– 12.

40- Societal attention toward extinction threats Ivan JaricCeline BellardFranck CourchampGregor KalinkatYves MeinardDavid L.RobertsRicardo A. Correia
Sci Rep 10, 11085 (2020).

Climate change had a significantly higher prominence than biological invasions within amphibians, birds and mammals, while the differences in the representation of the two threats within reptiles were not significant.

39Leclerc, CVilléger, SMarino, CBellard, C. Global changes threaten functional and taxonomic diversity of insular species worldwideDivers Distrib202026402– 414


38 -Englund, G., Öhlund, G., Olajos, F., Finstad, A., Bellard, C. and Hugueny, B. (2020), Holocene extinctions of a top predator – Effects of time, habitat area and habitat subdivision. J Anim Ecol. Accepted doi:10.1111/1365-2656.13174

37 –Li, X. ,H. Zhu, S. Geisen, C. Bellard, F. Hu, H. Li, X. Chen, M. Liu (2019), Agriculture erases climate constraints on soil nematode communities across large spatial scales. Global Change Biology. accepted 

36 -Li, X. , Bellard, C. , Hu, F. and Li, H. (2020). Effect of distance, area, and climate on the frequency of introduction and extinction events on islands and archipelagos. Ecosphere 11(1):e03008. 10.1002/ecs2.3008 pdf

35Li, X. , Bellard, C. , Hu, F. and Li, H. (2019), A comprehensive formula for decomposing change in community similarity into introduction and extinction events. Ecography. Accepted doi:10.1111/ecog.04613

34- Bellard, C. , Englund, G. and Hugueny, B. (2019), Biotic and abiotic drivers of species loss rate in isolated lakes. J Anim Ecol. doi:10.1111/1365-2656.12980


Extinction rate decreases with age since isolation of the fragments

33– T. Blackburn, Bellard, C. & T. Ricciardi, (2018) Alien versus native species as drivers of recent extinctions, in Frontiers in Ecology & Environment pdf

32– I. Castaneda, C. Bellard, I. Jaric, B. Pisanu, JL Chapuis & E. Bonnaud Global assessment of trophic patterns and home range size of red foxes (Vulpes vulpes) and domestic cats (Felis silvestris catus) in urban and peri-urban habitats, in Journal of Zoology pdf

31– C. Bellard, Jeschke J., Leroy B. & G. Mace, (2018) How climate change affects invasive species: insights from modelling studies across the globe, in Ecology & Evolution,, pdf 

30– C. Bellard*, J-F Rysman*, B. Leroy, C. Claud & G.M. Mace (2017) A global picture of biological invasion threat on islands, in Nature Ecology and Evolution doi:10.1038/s41559-017-0365-6 pdf


*equal contribution

29-Courchamp, A. Fournier, C. Bellard, C. Bertelsmeier, E. Bonnaud, J. M. Jeschke & J. Russel, Invasion Biology: specific problems and possible solutions in Trends in Ecology and Evolution pdf

28-Bradshaw C., Leroy B., Bellard C., Roiz D., Albert C., Barbet-Massin M., Fournier A., Salles JM, Simard F. & F Courchamp Massive yet grossly underestimated global costs of invasive insects, (2016) Nature communications  pdf

27-Hoffmann B., Luque G.M., Bellard C., Holmes N. & C. J. Donlan, A Global Review of Invasive Ant Eradications, Biological Conservation (2016) pdf

26Bellard, P. Genovesi & J. Jeschke, Global patterns in vertebrates threatened by biological invasions, Proceedings B 283(20152454) pdf

research highlight in Nature

25Bellard, P. Cassey & T. Blackburn, Alien species as a driver of recent extinctions, in Biology letters 12(20150623) pdf

research highlight in New Scientist

24Bellard & J. Jeschke A spatial mismatch between invader impacts and research activity, (2015) in Conservation biology 30, 230–232. pdf

Capture d_écran 2017-06-13 à 22.20.40

23-Gallardo, A. Ziertitz, T. Adriaens, C. Bellard, P. Boets, R. Britton, J. Newman, J. Valkenburg & D. Aldridge, Transnational horizon scanning for invasive non-native species: a cause study in western Europe, in Biological invasions 18, 17-30

22Bellard, B. Leroy, W. Thuiller, J-F Rysman & F. Courchamp, (2015) Major drivers of invasion risks throughout the world, Ecosphere, 7(3) pdf


21-G.M. Luque, C. Bellard, C. Bertelsmeier, E. Bonnaud, P. Genovesi, D. Simberloff, & F. Courchamp (2013) The 100th of the world’s worst invasive alien species. Biological invasion, 16, 981-985 pdf

20-G.M. Luque, C. Bellard, C. Bertelsmeier, E. Bonnaud, P. Genovesi, D. Simberloff & F. Courchamp. (2013) Monster fern makes IUCN invader list. Correspondence in Nature, 498 : 37 pdf

Climate and global changes 

19– B. Leroy, R.Delsol, B. Hugueny, C. Meynard, C. Barhoumi, M. Barbet-Massin & C. Bellard (2018) Without quality presence–absence data, discrimination metrics such as TSS can be misleading measures of model performance, in Journal of Biogeography, pdf

18– C. Leclerc, F. Courchamp & C. Bellard (2017) Insular threat association worldwide within islands and taxa, in Scientific reports, 8, doi:10.1038/s41598-018-24733-0 pdf


17Bellard, C. Leclerc & F. Courchamp, (2015) Combined impacts of global changes on biodiversity across the USA, in Scientific reports, 5, 11828 pdf

16Bellard, C. Leclerc, B. Hoffman & F. Courchamp, (2015) Vulnerability to climate change and sea-level rise of the 35th biodiversity hotspot, the Forests of East Australia, Environmental Conservation 43, 79-89 pdf

15Bellard, C*. Bertelsmeier C.*, P. Leadley, W. Thuiller & F. Courchamp (2012) Impacts of climate change on the future of biodiversity. Ecology letters, 15, 365-377*equal contribution pdf                   


14-Leroy, C. Bellard, A. Colliot, M. Vasseur, C. Courtial, M. Bakkenes, A. Carnard & F. Ysnel (2014), Forecasted climate and land use changes, and protected areas: the contrasting case of spiders, Diversity and Distribution, 20, 686-697 pdf

13-Marques, H.M. Pereira, C. Krug, P.W. Leadley, P. Visconti, S.R. Januchowski-Hartley, R.M. Krug, R. Alkemade, C. Bellard, W. L. Cheung, V. Christensen, H. D. Cooper, T. Hirsch, R. Hoft, J. van Kolck, T. Newbold, J. Noonan-Mooney, E. C. Regan, C. Rondinini, U. R. Sumaila, L. Teh & M. Walpole, A framework to identify enabling and urgent actions for the 2020 Aichi Targets, in Basic and Applied Ecology, 15, 633-638 pdf

12-P. Tittensor, M. Walpole, S. L. L. Hill, D. G. Boyce, G. L. Britten, Neil D. Burgess, S.H. M. Butchart, P. W. Leadley, E.C. Regan, R. Alkemade, R. Baumung, C. Bellard, et al., (2014), A mid term analysis of progress towards international biodiversity targets, Science, 346, 241-244 pdf

Biological invasions & Climate change

11-Gonzàlez-Munoz, C. Bellard, C. Leclerc, J-Y Meyer & F. Courchamp. (2015) Assessing current and future risks of invasion by Miconia calvescens D.C. Biological invasions 17, 3337-3350 pdf

10-Bellard, W. Thuiller, B. Leroy, M. Bakkeness, P. Genovesi & F. Courchamp. (2013) Will climate change promote future invasions? Global Change Biology, 19, 3740–3748 pdf


9Bellard, C. Leclerc, B. Leroy, M. Bakkeness, S. Veloz, W. Thuiller & F. Courchamp, Vulnerability of biodiversity hotspots to global changes, Global Ecology and Biogeography, 23, 1376-1386 pdf

Sea level rise

8Bellard, C. Leclerc & F. Courchamp. (2013) Impact of sea level rise on the ten insular biodiversity hotspots. Global Ecology and Biogeography, 23, 203-212 pdf

7Bellard, J. Russel, B. Hoffman, C. Leclerc & F. Courchamp (2014), Adapting island conservation to climate change. Response to Andréfouët et al. Trends in Ecology and Evolution, 30, 2-3 pdf

6-Courchamp, B. Hoffman, J. Russel, C. Leclerc & C. Bellard (2014) Keeping island afloat: climate change, sea-level rise and conservation, Trends in Ecology and Evolution, 29, 127-130 pdf

5Bellard, C. Leclerc & F. Courchamp (2013) Impact of sea level rise on the French islands. Nature Conservation 5: 75-86.

Other topics 

4-Leroy, C. Meynard, C. Bellard & F. Courchamp, (2015) Virtual species, an R package to generate virtual species distribution, Ecography 39, 599-607 pdf

3Leclerc*, C. Bellard*, G.M. Luque & F. Courchamp, Overcoming extinction: recovery of the doomed Tibetan antelope, Ecosphere, 6(9) *equal contribution

2-Cornille, T. Giraud, C. Bellard, A. Tellier, B. Le Cam, M. J. M Smulders, J. Kleinschmit, I. Roldan-Ruiz & P. Gladieux (2013) Postglacial recolonization history of the European crabapple (Malus sylvestris Mill.), a wild contributor to the domesticated apple. Molecular Ecology, 22, 2249-2263 pdf

1-Milner-Gulland E. J., Arroyo, B., Bellard, C., Blanchard, J., Bunnefeld, N., Delibes- Mateos, M., Edwards, C., Nuno, A., Palazy, L., Reljic, S., Riera, P., T. Skrbinsek (2010) New directions in Management Strategy Evaluation through cross-fertilisation between fisheries science and terrestrial conservation. Biology letters, 719-722 pdf

Authorship in expertise reports      

IPCC: Contributing Author of the Intergovernmental Panel on Climate Change, Working Group 2 Invasive species Chapter.

Settele J, Scholes R, Betts R, Bunn S, Leadley P, Nepstad D, Overpeck JT, Taboada MA (2014). Terrestrial and Inland Water Systems. In: Climate Change 2014: Impacts, Adaptation, and Vulnerability. Part A: Global and Sectoral Aspects. Contribution of Working Group II to the Fifth Assessment Report of the Intergovernmental Panel on Climate Change. Cambridge University Press, Cambridge, United Kingdom and New York, NY, USA, pp. 271-359. pdf

GBO-4: Lead Author of the Global Biodiversity Outlook-4 (Target 9: Invasive species)

Leadley, P., Krug, C., Alkemade, R., Pereira, H., Sumaila, U., Walpole, M., Marques, A., Newbold, T, Teh, L., van Kolck, J., Bellard, C., Januchowski-Hartley, S. and Mumby, P. (2014) Progress towards the Aichi Biodiversity Targets: an assessment of biodiversity trends, policy scenarios and key actions Montreal, QC, Canada: Secretariat of the Convention on Biological Diversity pdf

Articles under review : 

Leclerc C, F. Courchamp & C. Bellard Future climate change vulnerability of islands

Bellard, C.; Hugueny, B Importance of metapopulation dynamic to explain fish persistence in a river system, in minor revision Freshwater biology

Jaric, I.; Bellard, C.; Courchamp, F.; Kalinkat, G.; Meinard, Y.; Roberts, D.; Correia, R; Societal attention toward extinction threats bioRxiv 706911 i

Jaric, I.; Bellard, C.; Correia, R.; Courchamp, F.; Douda, K.; Essl, F.; Jeschke, J.; Kalinkat, G.; Kalous, L.; Lennox, R.; Novoa, A.; Proulx, R.; Pyšek, P.; Soriano-Redondo, A.; S., A.; Vardi, R.; Verissimo, D.; Roll, U. Invasion culturomics and iEcology approaches to better understand biological invasions